Brown spots on holly fern leaf undersides? Don't panic—those are sori (spore cases), a sign of a healthy, mature fern!
If fronds are pale or stunted, it's likely cultural: insufficient water, poor drainage, or nutrient deficiency. Holly ferns (Cyrtomium falcatum) thrive in Savannah with adequate moisture.

fern sori

American Holly Losing Leaves in Spring? It's Normal—They're Semi-Evergreen
Blister Beetles on Holly: Temporary Defoliators
Holly Scale Infestation: Identification and Treatment Plan
Resurrection Fern on Trees: Harmless Epiphyte
Liriope Anthracnose: Red-Brown Spots Mean Fungal Disease
Peace Lily Scale Infestation: To Treat or Toss?
Reviving Dormant Shamrocks: Summer Care for Healthier Growth
Distinguishing Environmental Leaf Spots from Disease on Wax Begonias
Troubleshooting Swamp Milkweed: Spider Mites, Aphids, and Webbing
Bougainvillea Cream Spots and Leaf Drop: Magnesium or Water
Holly Leaf Spots and Curling: Fungal Leaf Spot, Not Aphids
Oak Tree Webbed Clumps: Fall Webworms, Minor Pest
Paperwhite Bulbs: Black Streaks and Flopping—Light or Bacteria?
Troubleshooting Gloxinia Decline: Questions to Pinpoint the Cause
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