Some species—most famously the saguaro cactus (Carnegiea gigantea)—are sensitive to sudden changes in sun exposure. The side that faced north in the nursery (shaded) can sunscald if rotated to face south after transplant, leading to tissue death and pathogen entry.
Growers now spray-paint an "X" or mark the north side of the trunk on sensitive species. Installers align the mark to north in the landscape.
Thin-barked trees (maples, beeches, young fruit trees) can also benefit from orientation matching, though the effect is less dramatic than for saguaros.
If your spec requires orientation, it's likely for a species known to be sensitive. Follow it—cost is near zero, risk of non-compliance is real.
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