Rust appears as orange, brown, or reddish pustules on leaves and stems—caused by various fungal pathogens (primarily Puccinia and Uromyces species). While more common in humid climates, it can affect succulents even in dry areas if conditions allow.
Reference: eHow guide on detecting and preventing rust (www.ehow.com/how_3036_detect-prevent-rust.html)
Note: For cacti, avoid excessive handling of spines; treat only if rust is confirmed and cultural changes don't resolve it. Always test any product on a small area first.
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